Hugely popular Mozilla community providing support for Firefox, Thunderbird, Seamonkey and other well known Mozilla products. Over 350k members and 4 Million posts.
Inspired by Max Berry's book, Jennifer Government, NationStates is one of the most popular online simulation games. The forum boasts over 5 million posts and 200,000 users.
The VideoLAN project produces the extremely popular VLC media player software. Their forums provide support for users of the software as well as the development community.
OpenOffice.org offers a free and fully-featured office suite which meets the needs of the common consumer all the way up to large enterprises.
KDE is mainly known for producing the KDE Desktop Environment (used in Kubuntu) for Linux.
VirtualBox, developed by Oracle, is cross-platform x86 virtualisation software. It has become one of the most widely used virtualisation solutions in the world.