The VideoLAN project produces the extremely popular VLC media player software. Their forums provide support for users of the software as well as the development community.
Gentoo Linux is an open source operating system built on top of the Linux kernel and based on the Portage package management system.
Avant Browser is a fast and user friendly browser which uses Microsoft's Trident rendering engine. The browser has been downloaded over 22 million times.
Zend Technologies is one of the largest and most popular companies producing a wide variety of tools and frameworks for PHP. Their community forums rely on phpBB.
Discussion board for the massively popular xkcd web comic. Covering everything from Maths and Science to Logic Puzzles and Forum Games. Over 100k members and 2.4 Million posts.
OpenOffice.org offers a free and fully-featured office suite which meets the needs of the common consumer all the way up to large enterprises.