Gentoo Linux is an open source operating system built on top of the Linux kernel and based on the Portage package management system.
Discussion board for the massively popular xkcd web comic. Covering everything from Maths and Science to Logic Puzzles and Forum Games. Over 100k members and 2.4 Million posts.
OpenOffice.org offers a free and fully-featured office suite which meets the needs of the common consumer all the way up to large enterprises.
Pixelmator is a popular commercial image editing software for Mac OS X users.
Solia Online is one of the most active online communities with over 18 million posts. Solia Online allows you to create your own avatar, buy items, and discuss topics with your friends.
The VideoLAN project produces the extremely popular VLC media player software. Their forums provide support for users of the software as well as the development community.